It’s all there if you magnify the image!
In recent years, the Central American species A. cupreus has been split into six species. Jenny Gubler, working in Kirsten Nicholson’s lab at Central Michigan, presented a poster on the results of a genetic analysis of variation in this group. Examining mitochondrial DNA, she found support for the monophyly of the entire group and for existence of A. dariense, but that A. villai belongs within that species. Biogeographically, a clear separation is seen between those on the Pacific and Atlantic sides of the continental mountain range chains, and the clade is arrayed from north (basal) to south (highly nested), in support of previous work.
Professor of Biology and Director of the Living Earth Collaborative at Washington University in Saint Louis. I've spent my entire professional career studying anoles and have discovered that the more I learn about anoles, the more I realize I don't know.
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